Out-of-bounds sweeps consistently coincide with internal direction pointing towards the same location. Colour-coded 2D histograms of conditional occurrences of the two LMT latent variables (internal direction and sweeps in head-centred coordinates) for sweeps that terminate inside (top) or outside (bottom) visited portions of the environment in one animal (rat 25843, the same session as a – c ). The circular correlation coefficient between head-centred internal direction and sweep directions was similar when analysis was confined to theta cycles in which sweeps terminated inside versus outside the wagon-wheel track: r = 0.83 versus r = 0.82. Similar results were obtained for a second rat with fewer cells (not shown): r = 0.34 versus r = 0.35.
