Top, circular firing-rate plots showing broad tuning to head direction (HD) for four example cells in the parasubiculum. Middle, 2D position rate maps for the same cells. Bottom, temporal autocorrelograms of the cells firing rates. Arrows indicate temporal lags corresponding to the second theta peak (about 250 ms). Note the strong theta-cycle skipping in the first two cells, and conjunctive grid tuning in the last two.