Hippocampal sweeps follow MECparasubiculum sweeps. Top, decoded sweeps from co-recorded hippocampal cells (green) and MECparasubiculum cells (blue) over five successive theta cycles. Current location (arrowhead) and future path (grey line) are shown. Note the alignment of hippocampal and MECparasubiculum sweeps. Bottom, progression of decoded sweeps along the head direction axis as a function of time from the beginning of the theta cycle (for an example recording session). Note that hippocampal sweeps are delayed relative to MEC sweeps.