Sum of weights of place cells (black) and tone cells (pink) averaged by the time bin of peak activation of each assembly against the first lick. Assemblies active early in a trial were place cell dominated (black line). Assemblies preceding the lick (vertical black line) were tone cell dominated (pink line; n = 33 sessions from 6 mice; Wilcoxon two-sided signed-rank test, maximum P value in each block = 0.0273, 0.0332, 0.0234, 2.4 x 10 -4 , 0.0371, 4.185 x 10 -4 and 0.0269). Grey dashed horizontal line, equal contribution of place and tone cells. * P < 0.05 and *** P < 0.001. Data are shown as mean ± s.e.m.