Three example CA1 pyramidal cells displaying tone (left), mixed (middle) and place (right) tuning. Trajectories (grey lines) by trial type (top; correct trials only are shown, and spikes are overlaid in red), and the average firing rates as heatmaps with respect to position (place map) and frequency (frequency map) for no-tone and tone trials (bottom) are shown. The rainbow spectrum refers to trial types for ports 1–6.