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Refinement example for a filter whose weights had been initialized with the JASPAR profile of TFAP2C (MA0815.1). Freezing the filter weights during training was detrimental (top): the importance scores of that filter’s unit across all immune lineages were null. However, if filter weight refinement was allowed during training (i.e., no freezing; bottom), that filter was modified until it resembled the motif of EBF1, an important TF for maintaining B cell identity [ 51 ], and that same unit became important for predicting accessibility in B cells.
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