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Schematic showing that entorhinal cortex layer 3 (EC3) input to distal CA1 apical dendrites elicits mid-frequency gamma oscillations (red) at the peak of CA1 pyramidal layer theta (black trace), whereas CA3 inputs to proximal CA1 apical dendrites elicit slow gamma oscillations (blue) at the descending theta phase. Fast gamma oscillations (green) during theta troughs are generated by a local excitatory-inhibitory circuit motif.

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