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Phylogenetic tree of concatenated archaeal marker gene files was produced using GTDB-Tk (v.1.3.0) [ 29 ], with Methanomicrobium mobile BP used as the outgroup. Phylogeny was inferred using FastTree (v2.1.10) [ 30 ] and visualisation by iToL ( https://itol.embl.de/ ). MAGs and isolate genomes of 90% completeness and 5% contamination were included. Cultured isolates are identified by a red star. Bootstrap values are shown by the red ( 0.7) and black ( 0.9) circles. The black arrow denotes the common ancestor of Methanocorpusculum , the white arrow denotes the common ancestor of the Env clade and host clade 2, and the blue arrow denotes the common ancestor of the Env clade.

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