Individual voxels represent the lowest hierarchical resolution of our organelle, containing feature information such as voxel intensities and motility metrics. Shown here is a subset of motility features extracted for one tracked voxel (gray dots) through time T - 1 (blue structure) to T + 1 (red structure). The tracked reference point for each time point (blue dot, red dot) is shown, and is used to calculate reference-adjusted linear and angular movements.