Structure of Acs1 in its filamentous form. Left: helical array of one Acs1 filament. A single strand is highlighted in dark gray, and three consecutive layers are colored in different shades of blue. Right: enlarged view of the central three layers with the single helical strand indicated by a red arrowhead. The side and top view of the central Acs1 trimer are shown on the right. The 3-fold axis is represented by a triangle. See alsoFigures S1,S4, andS5.