Joint administration of EdU and tamoxifen 24 h before sacrifice identifies two-cell clones in the basal layer. Data are represented as mean SD.*p < 0.05,**p < 0.01***p < 0.001 Statistical tests are Wilcoxon matched-pairs signed ranked test (H) and two-sided paired t test (J). Scale bars: (H and K) 10 m and (D) 20 m. n = 47 in all experiments.