Top: snapshots of rotating doublets with labelled E-cadherin-mNG (green) and corresponding segmented meshes. Grey double arrows indicate the approximate elongation axis. Bottom left: a schematic for the definition of three doublet elongation axes a , b and c . Bottom right: relative doublet elongation magnitudes as a function of time after division (dots, individual data points; lines, connected binned averages). Inset: the ratio of elongation magnitudes, where positive values indicate a prolate shape.