Threatened species are those assessed as critically endangered, endangered or vulnerable (including those flagged as possibly extinct and possibly extinct in the wild). The following distributions are included: presence refers to extant, probably extant or possibly extinct; origin denotes native, reintroduced or assisted colonization; and seasonality indicates resident, breeding, non-breeding or passage. The value of each cell is calculated as the count of threatened species with a mapped distribution overlapping the cell. Richness is shown using a 0.5 x 0.5 latitude–longitude grid and WGS84. World Bank Official Boundaries (licensed under a Creative Commons licence CC BY 4.0 ) were used as the base map. For absolute threatened species richness of tetrapods, see ref. 33 .