Example beta-modulated cells for each region (one cell per row), with firing relative to beta-bout troughs (one trough per bout) in either PFC, NAc, Amy, or Hpc CA1 (one region beta-trough per column; 1.5-ms bins). Note that triggering neuronal spiking by beta troughs (separated by at least 250 ms) shows at least three peaks of rhythmic firing modulation matching beta frequency.